Swiss Journal of Geosciences

Table 9 Characterization of trackways corresponding to the four main locomotion styles of limulids

From: A giant limulid trackway (Kouphichnium lithographicum) from the lithographic limestones of Cerin (Late Kimmeridgian, France): ethological and environmental implications

Behaviour
Locomotion style	WALKING	CRAWLING	DYING	PLOUGHING
Locomotion rate	High	Medium	Low	Very low
Resulting traces
Trackway	Straight	Straight	Sinuous	Sinuous
Prosoma imprints	Absent	Absent	Slight	Deep
Telson imprints	Absent	Continuous	Interrupted	Interrupted
Pushers imprints	Well marked	Well marked	Slight	Absent
Walking legs imprints	Short	Short	Long	Absent
Series length	High	Medium	Low	X
Repeat distance	High	Medium	Low	X
Imprint spacing	High	Medium	Low	X
Angle to mid-line	Low	Medium	High	X

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